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The fossil record of beavers dates to the late Eocene of Asia, and beavers reached North America and Europe in the early Oligocene. Several lines of descent developed in the tertiary. One line developed fossorial adaptations; the fossil remains of the northern American miocene beaver Palaeocastor have been found in spectacular corkscrew shape burrows dug by these animals. Another evolutionary line led to the bear sized giant beaver castorids of the North American pleistocene. Throughout their history, castorids have been restricted to the northern hemisphere.
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