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A 52-year- old male sought medical help complaining about progressive appearance of hyperchromic (hyper pigmented) papules on the edge of the second finger of both hands. He also complained about darkening of urine. Incisional biopsy of one of the spots on his hands and urine sample tested positive for or homogentisic acid. Based on these findings a diagnosis of alkaptonuria is made. This diagnosis is consistent with a deficiency in which of the following enzymes?
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